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This is the more advanced Diptera suborder, which includes 84 families separated into various clades or unranked divisions, infraorders and superfamilies. The primary distinguishing characteristic is that unlike the Nematocera species that have long segmented antennae, all the Brachyceran flies have reduced antennal segments, as the flagella are fused together with the remaining diminutive segments forming a stumpy style or bristle-like arista. Whereas Nematocera effectively means species with ‘thread-horns’, Brachycera means ‘shortened-horns’.

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The following listing provides a personal reference document of all the species I've seen and have managed to photograph but, whereas the thumbnail images for most of the species in other orders link to a dedicated 'species account', there are simply too many individual Diptera species, so the arrangement here is different. Clicking any of the thumbnail images below will take you to a common 'family account' page, which includes a bit of basic information together with photos of each species. The details generally relate to the family as a whole but, for some species, there are extra notes with features to assist identification.

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LOWER BRACHYCERA

An unranked division of 11 families that are currently separated into five superfamilies and four infraorders, which together form what is almost certainly the most well-covered group of families. With the exception of the Syrphidae (Hoverfly) species that are equally well documented, all of my 'family accounts' in the other divisions quote 'general characters' as supportive information is sparse. That's certainly not the case with these families as they are all extensively covered within the excellent, and highly recommended, illustrated identification guide entitled 'British Soldierflies and their Allies' by Alan Stubbs and Martin Drake. The group is unusual in that all of the species have a recognised common name. Although I have a long way to go with DIPTERA species in general, I have photographed at least a couple of species in each of the 6 major families of this division. The remaining 5 families are all very small with only two or three members so may not feature even in the future. 

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Lower Brachycera

Rhagionidae (Snipeflies)

Chrysopilus asiliformis

Chrysopilus asilliformis

Little Snipefly

Rhagio lineola

Rhagio lineola

Small Flecked-winged Snipefly

Tabanidae (Horseflies)

Atylotus latistriatus

Atylotus latistriatus

Saltmarsh Horsefly

Tabanus bromius

Tabanus bromius

Band-eyed Brown Horsefly

Chrysopilus cristatus

Chrysopilus cristatus

Black Snipefly

Rhagio scolopaceus

Rhagio scolopaceus

Downlooker Snipefly

Rhagio tringarius

Rhagio tringarius

Marsh Snipefly

Haematopota pluvialis

Haematopota pluvialis

Notch-horned Cleg

Hybomitra distinquenda

Hybomitra distinquenda

Bright Horsefly

Tabanus sudeticus

Tabanus sudeticus

Giant Horsefly

Stratiomyidae (Soldierflies)

Beris geniculata

Long-horned Black Legionnaire

Beris geniculata
Chloromyia formosa

Chloromyia formosa

Broad Centurion

Microchrysa polita

Microchrysa polita

Black-horned Gem

Oplodontha viridula

Oplodontha viridula

Common Green Colonel

Bombyliidae (Bee-flies)

Bombylius discolor

Bombylius discolor

Dotted Bee-fly

Thyridanthrax fenestratus

Thyridanthrax fenestratus

Mottled Bee-fly

Bombylius major

Bombylius major

Dark-edged Bee-fly

Bombylius minor

Bombylius minor

Heath Bee-fly

Villa cingulata

Villa cingulata

Downland Villa

Therevidae (Stiletto-flies)

Thereva nobilitata

Thereva nobilitata

Common Stiletto

Thereva plebeja

Thereva plebeja

Crochet-hooked Stiletto

Asilidae (Robberflies)

Asilus crabroniformis

Asilus craboniformis

Hornet Robberfly

Choerades marginatus

Choerades marginatus

Golden-haired Robberfly

Machimus atricapillus

Machimus atricapillus

Kite-tailed Robberfly

Machimus cingulatus

Machimus cingulatus

Brown Heath Robberfly

Dioctria baumhaueri

Dioctria baumhaueri

Stripe-legged Robberfly

Dioctria linearis

Dioctria linearis

Small Yellow-legged Robberfly

Dioctria rufipes

Dioctria rufipes

Common Red-legged Robberfly

Dysmachus trigonus

Dysmachus trigonus

Fan-bristled Robberfly

 

EREMONEURA

An intermediate unranked clade of BRACHYCERA that in some classification systems includes the superfamily Empidoidea as well as the Cyclorrhapha (see below), which was once regarded as a third suborder. Current taxonomy lists 5 families that were traditionally referred to as the ‘Dolis and Empids’, the ‘Dolis’ for the Long-legged Flies of the Dolichopodidae family, and the ‘Empids’ for the Dance-flies that were all formally within Empididae, which is now split into separate families.

 

   

Eremoneura

Hybotidae (Dance-flies)

Bicellaria sp.

Bicellaria sp.

Hybos culiciformis

Hybos culiciformis

Platypalpus sp.

Platypalpus sp.

Tachydromia arrogans

Tachydromia arrogans

Empididae (Dagger-flies)

Empis albinervis

Empis albinervis

Empis tessalata

Empis tessalata

Empis livida

Empis livida

Empis trigramma

Empis trigramma

Empis lutea

Empis lutea

Empis pennipes

Empis pennipes

Hilara cf.maura

Hilara cf.maura

Dolichopodidae (Long-legged Flies)

(now split across two pages to provide a separate photo gallery for a number of unidentified species)

Dolichopus sp.

Dolichopus sp.

Dolichopus cf.griseipennis

Dolichopus cf.griseipennis

Dolichopus cf.ungulatus

Dolichopus cf.ungulatus

Dolichopodidae sp.

unidentified species

Medetera cf.diadema

Medetera cf.diadema

Neurigona quadrifasciata

Neurigona quadrifasciata

Poecilobothrus nobilitatus

Poecilobothrus nobilitatus

Sciapus platypterus

Sciapus platypterus

Campsicnemus scambus

Campsicnemus scambus

Hydrophorus cf.oceanus

Hydrophorus cf.oceanus

Aschiza

 

MUSCOMORPHA

The largest division of BRACHYCERA; an infraorder that in most current classification systems is being used to replace Cyclorrhapha. Although, to all intents and purposes the two terms are synonymous, it's worth remembering that Cyclorrhapha relates to the manner in which these flies exit their puparium, which is through a special end-cap within a weakened circular seam (cyclor meaning circular, and rhapha meaning seam). Most new adults aid their emergence by inflating part of their face, the ptilinum, which sinks back after they exit to leave a sunken area surrounded by a horseshoe-shaped groove known as the ptilinal suture. All the flies that use this method are known as the Schizophora. Some of the flies that are closer to the LOWER BRACHYCERA lack this ability and thus have no ptilinal suture and are separated into a relatively small subdivision called the Aschiza.

 

Aschiza

This subdivision includes just 6 families, one of which is Syrphidae. Of all the DIPTERA families, no other group generates more interest, or has a greater following, than Hoverflies, which is why the thumbnails of the representative species below links to a separate dedicated page for the species that I've been fortunate enough to see and photograph.     

 

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Platypezidae (Flat-footed Flies)

Protoclythia modesta (male)

Introduction, start here

Agathomyia falleni

Agathomyia falleni

Agathomyia unicolor

Agathomyia unicolor

(an extended section with separate pages for the two main subfamilies Callomyiinae and Platypezinae)

Platypeza consobrina

Platypeza consobrina

Platypeza sp.

Platypeza sp.

Protoclythia modesta

Protoclythia modesta

Protoclythia rufa

Protoclythia rufa

Phoridae (Scuttleflies)

Phora sp.

Phora sp.

Triphleba minuta

Triphleba minuta

Triphleba papillata

Triphleba papillata

Lonchopteridae (Spear-winged Flies)

Lonchoptera cf.lutea

Lonchoptera cf.lutea

Syrphidae (Hoverflies)

(as previously noted, clicking any of the following thumbnails will take you to a separate page currently featuring 70+ species)

Chrysotoxum festivum

Chrysotoxum festivum

- representative species -

Episyrphus balteatus

Episyrphus balteatus

- representative species -

Riponnensia splendens

Riponnensia splendens

- representative species -

Volucella zonaria

Volucella zonaria

- representative species -

Acalyptrates

 

Schizophora

This is the major subdivision of MUSCOMORPHA consisting 62 families that are separated into two further unranked sections, called Acalyptratae and Calyptratae, more commonly known as the Acalyptrates and the Calyptrates.

 

Acalyptrates

Whilst Schizophora is by far the larger of the two divisions of MUSCOMORPHA, the Acalyptratae is the single largest section of the BRACHYCERA suborder representing no less than 49 families and around 1600 species. Given that the Acalyptrates are such a diverse group of species it's not surprising to note that the families are separated into ten superfamilies. Each of the accompanying 'family accounts' confirms the higher taxonomic structure for that particular family. The Acalyptrate species are defined by their lack of calypters and certain other features that characterise the Calyptrates (as explained below). 

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Micropezidae (Stilt-legged Flies)

Neria cibaria

Neria cibaria

Megamerinidae (Bearded Fool Fly)

Megamerina dolium

Megamerina dolium

Psilidae (Rust Flies)

Imantimyia albiseta

Imantimyia albiseta

Psila fimetaria

Psila fimetaria

Conopidae (Thick-headed Flies)

Conops quadrifasciatus

Conops quadrifasciatus

Myopa pellucida

Myopa pellucida

Myopa sp.

Myopa sp.

Sicus ferrugineus-6.jpg

Sicus ferrugineus

Lonchaeidae (Lance Flies)

Lonchaeidae sp.

Lonchaeidae sp.

Ulidiidae (Picture-winged Flies)

Seioptera vibrans.jpg

Seioptera vibrans

Platystomatidae (Signal or Semaphore Flies)

Platystoma seminationis

Platystoma seminationis

Tephritidae (True Fruit Flies)

Anomoia purmunda

Anomoia purmunda

Chetostoma curvinerve

Chetostoma curvinerve

Tephritis formosa

Tephritis formosa

Tephritis neesii

Tephritis neesii

Lauxaniidae (Lauxaniid Flies)

Calliopum cf.simillium

Calliopum cf.simillium

Meiosimyza rorida

Meiosimyza rorida

Minettia inusta

Minettia inusta

Homoneura notata

Homoneura notata

Sapromyza sexpunctata

Sapromyza sexpunctata

Sapromyzosoma quadripunctata

Sapromyzosoma quadripunctata 

Euleia heraclei

Euleia heraclei  (black morph)

Urophora cardui

Urophora cardui

Euleia heraclei

Euleia heraclei  (brown morph)

Urophora jaceana

Urophora jaceana

Minettia longipennis

Minettia longipennis

Tricholauxania cf.praeusta

Tricholauxania cf.praeusta

Lauxaniidae sp.

Lauxaniidae sp.

Coelopidae (Kelp or Wrack Flies)

(includes a separate page for comparing various 'seaweed flies' that are featured in other families)

Coelopa frigida

Coelopa frigida

Coelopa pilipes

Coelopa pilipes

Malacomyia sciomyzina

Malacomyia sciomyzina

Dryomyzidae (Dryomyzid Flies)

Dryomyza anilis

Dryomyza anilis

Heterocheilidae (Oarweed Fly)

Heterocheila buccata

Heterocheila buccata

Sciomyzidae (Marsh Flies)

Coremacera marginata

Coremacera marginata

Elgiva cucularia

Elgiva cucularia

Renocera pallida

Renocera pallida

Pherbina coryleti

Pherbina coryleti

Sepsidae (Black Scavenger Flies)

Pherbellia dubia

Pherbellia dubia

Tetanocera ferruginea

Tetanocera ferruginea

Pherbellia ventralis

Pherbellia ventralis

Trypetoptera punctulata

Trypetoptera punctulata

Orygma luctuosum

Orygma luctuosum

Sepsis cynipsea.jpg

Sepsis cynipsea

Sepsis sp.jpg

Sepsis sp.

Clusiidae (Clusiid or Druid Flies)

Clusiodes verticalis

Clusiodes verticalis

Agromyzidae (Leaf-mining Flies)

Phytomyza ranunculi-2.jpg

Phytomyza ranunculi

Opomyzidae (Opomyzid Flies)

Geomyza cf.balachowskyi

Geomyza cf.balachowskyi

Geomyza tripunctata

Geomyza tripunctata

Opomyza florum

Opomyza florum

Opomyza germinationis

Opomyza germinationis

Milichiidae (Freeloader Flies)

Milichiidae sp.

Milichiidae sp.

Chloropidae (Grass Flies)

Chlorops cf.hypostigma

Chlorops cf.hypostigma

Thaumatomyia cf.hallandica

Thaumatomyia cf.hallandica

Chlorops cf.pumilionis

Chlorops cf.pumilionis

Thaumatomyia notata

Thaumatomyia notata

Chlorops sp.

Chlorops sp.

Lasiambia sp.

Lasiambia sp.

Heleomyzidae (Spiny-winged Flies)

Suillia affinis

Suillia affinis

Suillia sp.

Suillia sp.

Suillia atricornis

Suillia atricornis

Heleomyzidae sp.

Heleomyzidae sp.

Suillia bicolor

Suillia bicolor

Suillia variegata

Suillia variegata

Tephrochlamys rufiventris

Tephrochlamys rufiventris

Sphaeroceridae (Lesser Dungflies)

Copromyza equina

Copromyza equina

Copromyzinae sp.

Copromyzinae sp.

Copromyza nigrina

Copromyza nigrina

Crumomyia fimetaria

Crumomyia fimetaria

Crumomyia roserii

Crumomyia roserii

Rachispoda sp.

Rachispoda sp.

Drosophilidae (Small Fruit Flies)

Drosophilia melanogaster

Drosophila melanogaster

Drosophila cf.suzukii

Drosophila cf.suzukii

Drosophilidae - undetermined species

Drosophila sp.

Diastatidae (Diastatid Flies)

Diastata fuscula

Diastata fuscula

Ephydridae (Shore Flies)

(this family account now separates the freshwater / wetland species from those found in coastal and saltmarsh areas)

Notiphila cf.riparia

Notiphila cf.riparia

Notiphila sp.2

Notiphila sp.2

Notiphila sp.3

Notiphilia sp.3

Dichaeta caudata

Notiphila (Dichaeta) caudata

Ochthera mantis

Ochthera mantis

Parydra coarctata

Parydra coarctata

Parydra littoralis

Parydra littoralis

Ephydra sp.

Ephydra sp.

Paracoenia fumosa

Paracoenia fumosa

Calyptrates

 

Calyptrates

This final section includes flies with usually large and well developed calypters (except in Scathophagidae), often concealing the halteres when viewed from above; a thorax with well-defined greater ampulla; scutum with complete transverse suture and strongly developed posterior calli; and cleft in the second antennal segment.

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There are 13 families separated into three superfamilies; Hippoboscoidea, which is not currently represented here, consisting the Hippoboscidae louse fly and Nycteribiidae bat fly families; then Muscoidea and Oestroidea, which include a number of familiar species, such as the common yellow dungfly, housefly, stable fly and the generically termed 'bluebottle' and 'greenbottle' blowflies.

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Scathophagidae (Dungflies)

(this family account has now been split to allow the Scathophaga 'true dungfly' species to be featured on a separate page)

Scathophaga cf.inquinata

Scathophaga cf.inquinata

Cordilura ciliata

Cordilura ciliata

Scathophaga furcata

Scathophaga furcata

Norellisoma spinimanum

Norellisoma spinimanum

Scathophaga cf.scybalaria

Scathophaga cf.scybalaria

Scathophaga stercoraria

Scathophaga stercoraria

Ceratinostoma ostiorum

Ceratinostoma ostiorum

Scathophaga suilla

Scathophaga suilla

Scathophaga litorea

Scathophaga litorea

Anthomyiidae (Root-maggot flies)

(an extended section that is now broadly separated by genera and spread across six pages) 

unidentified anthomyiid

Introduction, start here >

Emmesomyia cf.socia

Emmesomyia cf.socia

Anthomyia procellaris

Anthomyia procellaris

Hylemya cf.vagans

Hylemya cf.vagrans

Hydrophoria cf.lancifer

Hydrophoria cf.lancifer

unidentified anthomyiid

unidentified anthomyiid

Anthomyia pluvialis (complex)

Anthomyia pluvialis (complex)

Pegomya cf.bicolor

Pegomya cf.bicolor

Hylemya cf.variata

Hylemya cf.variata

Hydrophoria cf.linogrisea

Hydrophoria cf.linogrisea

Unidentified anthomyiid

unidentified anthomyiid

Unidentified anthomyiid

unidentified anthomyiid

Pegomya cf.geniculata

Pegomya cf.reticulata

Anthomyia cf.imbrida

Anthomyia cf.imbrida

Hylemya vagans-nigrimana

Hylemya vagrans /nigrimana

Hydrophoria cf.ruralis

Hydrophoria cf.ruralis

Coastal and strandline species .....

Fucellia tergina-1.jpg

Fucellia tergina

Unidentified strandline anthomyiid

unidentified anthomyiid

Unidentified strandline anthomyiid

unidentified anthomyiid

Unidentified strandline anthomyiid

unidentified anthomyiid

Muscidae (Houseflies)

(an extended section with species broadly grouped in taxonomic order by subfamily and featured across seven pages) 

Graphomya maculata

Introduction, start here >

Coenosia tigrina

Coenosia tigrina

Lispe tentaculata

Lispe tentaculata

Eudasyphora cyanella

Eudasyphora cyanella

Morellia hortorum

Morelia hortorum

Polietes cf.lardarius

Polietes cf.lardarius

Graphomya maculata

Graphomya maculata

Mydaea corni

Mydaea corni

Phaonia angelicae

Phaonia angelicae

Helina evecta

Helina evecta

Phaonia tuguriorum

Phaonia tuguriorum

Hydrotaea cf.dentipes

Hydrotaea cf.dentipes

Schoenomyza litorella

Schoenomyza litorella

Eudasyphora cyanicolor

Eudasyphora cyanicolor

Lispe cf.caesia

Lispe cf.caesia

Musca autumnalis

Musca autumnalis

Thricops diaphanus

Thricops diaphanus

Limnophora tigrina

Limnophora tigrina

Lispe cf.nana

Lispe cf.nana

Mesembrina meridiana

Mesembrina meridiana

Neomyia cornicina

Neomyia cornicina

Muscina levida

Muscina levida

Limnophora triangula

Limnophora triangula

Neomyia viridescens

Neomyia viridescens

Stomoxys calcitrans

Stomoxys calcitrans

Hebecnema cf.vespertina

Hebecnema cf.vespertina

Mydaea cf.humeralis

Mydaea cf.humeralis

Helina cf.depuncta

Helina depuncta

Phaonia pallida

Phaonia pallida

Mydaea cf.urbana

Mydaea cf.urbana

Helina cf.impuncta

Helina impuncta

Phaonia rufiventris

Phaonia rufiventris

Helina sp-2.jpg

Helina sp.

Phaonia subventa

Phaonia subventa

Phaonia valida

Phaonia valida

Fanniidae (Lesser Houseflies)

Fannia sp.

Fannia sp.

Fannia cf.lustrator

Fannia cf.lustrator

Calliphoridae (Blowflies)

(this family has now been split to allow the Lucilia 'greenbottle' species to be featured on a separate page)

Calliphora vicina

Calliphora vicina

Melinda viridicyanea-3.jpg

Melinda viridicyanea

Lucilia cf.ampullacea-2.jpg

Lucilia cf.ampullaca

Lucilia cf.silvarum

Lucilia cf.silvarum

Polleniidae (Cluster Flies)

Pollenia cf.angustigena

Pollenia cf.angustigena

Calliphora vomitoria

Calliphora vomitoria

Melanomya nana

Melanomya nana

Lucilia cf.caesar

Lucilia cf.caesar

Calliphora sp.

Calliphora sp.

Protocalliphora azurea

Protocalliphora azurea

Lucilia cf.richardsi

Lucilia cf.richardsi

Lucilia cf.sericata

Lucilia cf.sericata

Pollenia cf.labialis

Pollenia cf.labialis

Pollenia cf.rudis

Pollenia cf.rudis

Pollenia sp.

Pollenia sp.

Rhinophoridae (Woodlouse Flies)

Rhinophora lepida

Rhinophora lepida

Sarcophagidae (Fleshflies)

Sarcophaga cf.haemorrhoa

Sarcophaga cf.haemorrhoa

Metopia cf.argyrocephala

Metopia cf.argyrocephala

Miltogramma germari

Miltogramma germari

Sarcophaga sp. (carnaria group)

Sarcophaga sp. (carnaria group)

Miltogramma punctata

Miltogramma punctata

Nyctia halterata

Nyctia halterata

Sarcophaga sp.

Sarcophaga sp.

Tachinidae (Tachinid Flies)

(the species in this extended two-page section are currently listed alphabetically, rather than in taxonomic order) 

Dexia rustica

Dexia rustica

Ectophasia crassipennis

Ectophasia crassipennis

Gymnocheta viridis

Gymnocheta viridis

Macquartia grisea

Macquartia grisea

Nowickia ferox

Nowickia ferox

Siphona sp.

Siphona sp.

Thelaria nigrina

Thelaira nigrina

Dexiosoma caninum

Dexiosoma caninum

Eriothrix rufomaculata

Eriothrix rufomaculata

Dinera grisescens

Dinera grisescens

Exorista rustica

Exorista rustica

Gymnosoma rotundatum

Gymnosoma rotundatum

Macquartia tenebricosa

Macquartia tenebricosa

Pales pavida

Pales pavida

Siphona cf.geniculata

Siphona cf.geniculata

Thelaria cf.solivaga

Thelaira cf.solivaga

Meigenia cf.mutabilis

Meigenia cf.mutabilis

Mintho rufiventris

Mintho rufiventris

Phania funesta

Phania funesta

Tachina fera

Tachina fera

Thelaria sp.

Thelaira sp.

Voria ruralis

Voria ruralis

Original list created Dec.21 | updated Dec.24 (v.11)

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